Forage fish play a crucial role in marine food webs in many ecosystems (Box 1.1). These small and medium-sized pelagic species are the primary food source for many marine mammals, seabirds, and larger fish, transferring energy from plankton to larger predators. Forage fish are also important predators in marine ecosystems, feeding upon phytoplankton, zooplankton, and, in some cases, the early life stages of their predators.
Forage fish play an intermediary role in many marine ecosystems, including estuaries, shelf seas, upwelling, and open ocean systems occurring from the tropics to the Earth’s poles. They constitute the majority of prey upon which some predators depend. Such highly dependent predators may be iconic or ecologically important, while others may be commercially or recreationally valuable fish species. In some cases, highly dependent predators may include threatened or endangered species. A reduction in available prey—because of fishing, environmental conditions, or a combination of both—can have direct and lasting impacts and can fundamentally change the structure and functioning of an ecosystem.